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This depends on how the application uses the JGroups channel. Referring to the example above, if there was something about the application that would naturally cause A to slow down its rate of sending because D wasn't keeping up, then FC would not be needed.
A good example of such an application is one that makes synchronous group RPC calls (typically using a JGroups RpcDispatcher.) By synchronous, we mean the thread that makes the call blocks waiting for responses from all the members of the group. In that kind of application, the threads on A that are making calls would block waiting for responses from D, thus naturally slowing the overall rate of calls.
A JBoss Cache cluster configured for REPL_SYNC is a good example of an application that makes synchronous group RPC calls. If a channel is only used for a cache configured for REPL_SYNC, we recommend you remove FC from its protocol stack.
And, of course, if your cluster only consists of two nodes, including FC in a TCP-based protocol stack is unnecessary. There is no group beyond the single peer-to-peer relationship, and TCP's internal flow control will handle that just fine.
Another case where FC may not be needed is for a channel used by a JBoss Cache configured for buddy replication and a single buddy. Such a channel will in many respects act like a two node cluster, where messages are only exchanged with one other node, the buddy. (There may be other messages related to data gravitation that go to all members, but in a properly engineered buddy replication use case these should be infrequent. But if you remove FC be sure to load test your application.)